Continuity mistake: In the season 5 episode 'Destiny' we see that the first time Angelus meets William, he has already adopted a more cockney accent, but in an earlier episode of BtVS (Fool For Love) Angelus asks Spike "Since when do you talk like that", suggesting he changed his accent after they met.
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Trivia: In the opening credits there is a shot of a girl standing alone on the corner of a street. This was taken from the Buffy episode "Anne", when Buffy was staying in Los Angeles.
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Answer: Due to his epiphany, Angel basically had the 'true darkness' he worked so hard to reattain ripped away from him, like his soul was after sleeping with Buffy.